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Aristida macroclada Henrard

Derivation
Aristida L., Sp. Pl. 1: 82 (1753); from the Latin arista (an awn).

macroclada- from the Latin macro (large) and clados (branch), derivation uncertain.

Published in
Meded. Rijks. Herb. Leiden 54A: 325 (1927).

Common synonyms
Aristida novaeguineae Ohwi

Habit
Perennial, tufted. Culms 60–100 cm tall, 5–9-noded. Mid-culm internodes smooth, glabrous. Lateral branches sparsely branched. Leaf-sheaths glabrous on surface. Ligule a fringe of hairs, 0.3–0.5 mm long. Collar glabrous. Leaf-blades straight, flat or conduplicate or involute or convolute, 15–30 cm long, 1–2.3 mm wide. Leaf-blade surface glabrous or pilose.

Inflorescence
Inflorescence compound, a panicle. Panicle open, elliptic, 17–24 cm long, 3–7 cm wide. Primary panicle branches appressed, moderately spaced, 5–10 cm long. Panicle branches with prominent pulvini.

Spikelets
Spikelets solitary. Fertile spikelets 1-flowered, comprising 1 fertile floret, without rhachilla extension, lanceolate, terete, 6–15 mm long, breaking up at maturity. Spikelets disarticulating below each fertile floret. Floret callus elongated, 0.5–0.8 mm long, pubescent, acute.

Glumes
Glumes persistent, similar, thinner than fertile lemma. Lower glume lanceolate, 4–11 mm long, 66–80% length of upper glume, membranous, 1-keeled, 1-nerved. Lower glume lateral nerves absent. Lower glume surface smooth to scabrous. Lower glume apex emarginate, mucronate. Upper glume lanceolate, 6–15 mm long, 110–160% of length of adjacent fertile lemma, membranous, 1-keeled, 1-nerved. Upper glume lateral nerves absent. Upper glume apex entire or erose, emarginate or cuspidate, mucronate.

Florets
Fertile lemma lanceolate, subterete, 5.5–9.5 mm long, coriaceous, 3-nerved. Lemma surface tuberculate or muricate, rough above. Lemma margins convolute, covering most of palea. Lemma apex acute, 3-awned. Median (principal) awn 3-branched, 11–37 mm long overall, 10–29 mm long, with a straight or slightly twisted column or with a twisted column, persistent. Column 1–8 mm long. Lateral lemma awns present, 9–26 mm long, shorter than principal, 90% of length of principal. Palea 20% of length of lemma, without keels. Anthers 3. Grain with adherent pericarp, terete, not grooved, 5–8 mm long. Embryo 25–50% of length of grain. Hilum linear.


Continental Distribution:
Tropical Asia, Australasia

Australian Distribution:
Northern Territory, Queensland.

Northern Territory: Darwin & Gulf. Queensland: Cook, North Kennedy.

Classification. (GPWG 2001):
Aristidoideae

Notes
Native. It exists as two subspecies, Aristida macroclada subsp. macroclada with a very short column and Aristida macroclada subsp. queenslandica with a fairly well developed column. Aristida macroclada differs from Aristida psammophila and Aristida longicollis by the inflorescence being loose to open and the branches being naked at the base.


Column of many spirals A. macroclada subsp. queenslandica (QLD)
Column of 1/2-1 spiral A. macroclada subsp. macroclada (NT QLD)
A. novaeguineae

Aristida macroclada Henrard subsp. macroclada
The "top end" of the Northern Territory, one record from Cape York Peninsula, QLD, and a few records from New Guinea. Eucalyptus, and Melaleuca communities on sandy and loamy soils. Flowering and fruiting January (one record), March to July.

Aristida macroclada Henrard subsp. queenslandica B.K.Simon
Tropical coastal Queensland. One record from New Guinea. Eucalyptus, Casuarina and Melaleuca communities on sandy and loamy soils. Flowering and fruiting February to June, August.


Images
Illustrations available:
Inflorescence (photo)
Spikelets, subsp. macroclada left, subsp. queenslandica right (line drawing)
Australian distribution



Inflorescence (photo)
© Queensland Herbarium
Sharp 423 and Simon
by D.Sharp


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Spikelets, subsp. macroclada left, subsp. queenslandica right (line drawing)
© Simon 1992
by Will Smith


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Australian Distribution
© ABRS


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