This project was supported by a grant from CANRI (Community Access to Natural Resource Information). Visit the CANRI website at

The authors thank the following Coleopterists who provided help or information, or whose work was instrumental in allowing us to construct this key: Mark Hanlon, Jack Hasenpusch, John Lawrence, Tom Weir and Geoff Williams.

Thanks to many of our colleagues who helped with advice, support, testing, encouragement and criticism, especially Gerry Cassis, Michael Elliott, Paul Flemons, Rebecca Harris, Tim Moulds and Kindi Smith.