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Chlorosis between veins on a mature leaf (J. O'Sullivan).  


Browning (necrosis) spreading from the margins  to areas between veins  (J. O'Sullivan).

Diagnostic summary

  • What you see on plants

- dark grey necrosis on old leaves spreading from margins into interveinal areas.


- dark necrotic spots on mature leaves, aligned midway between the main veins.


- necrosis is not preceded by yellowing, but affected leaves eventually yellow and fall off.


- crop wilts readily in dry weather.

  • Apart from its direct toxic effects, high salt levels can inhibit water uptake, and damage soil structure.

  • Occurs in coastal areas affected by sea water, or irrigated crops in semi-arid areas.

Characteristics and occurrence


Confusion with other symptoms

Diagnostic tests



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